Levings, C.D. 1968. Fertilized eggs of the butter sole, Isopetta isolepis in Skidegate Inlet, British Columbia. J. Fish. Res. Bd. Canada, 28(8): 1743 1744.

Levings, C.D. 1969. The zoarcid Lycodopis pacifica in outer Burrard Inlet, British Columbia. J. Fish. Res. Bd. Canada, 26(9): 2403 2412.

Levings, C.D. 1973. Dominance and movements of winter flounder (Pseudopleuronectes americanus) in St. Margaret’s Bay, Nova Scotia. Naturaliste Can., 100: 337 345.

Levings, C.D. 1974. Seasonal changes in feeding and particle selection by winter flounder (Pseudopleuronectes americanus). Trans. American Fish Society, 103: 828-832.

Levings, C.D. 1975. Analyses of temporal change in a shallow-water benthic community in Nova Scotia. Int. Revue ges. Hydro biol., 60(4): 449 470.

Hargrave, B.T., and C.D. Levings. 1975. A prognosis for benthic biological oceanography in Canada. In: T.R. Parsons, Review of Biological Oceanography in Canada. J. Fish. Res. Bd. Canada, 32(10): 1910 1961.

Levings, C.D. 1976. Basket traps for surveys of a gammarid amphipod (Anisogammarus confervicolus) at two British Columbia estuaries. J. Fish. Res. Bd. Canada 33: 2066 2069.

Levings, C.D. 1976. Reply to D.V. Ellis’ comments on a prognosis for Canadian benthic biological oceanography. J. Fish. Res. Bd. Canada 33: 1864.

Vermeer, K., and C.D. Levings. 1977. Populations, biomass, and food habits of ducks on the Fraser Delta intertidal area, British Columbia. Wildfowl 28: 49 60.

Wu, R.S.S., C.D. Levings, and D.J. Randall. 1977. Different strategies in energy partition between crowded and uncrowded individual barnacles (Balanus glandula Darwin). Can. J. Zool. 55: 642 647.

Chang, B.D., and C.D. Levings. 1978. Effects of burial on the heart cockle (Clinocardium nuttallii) and the Dungeness crab (Cancer magister). Estuarine and Coastal Marine Science, 7: 409 412.

Levings, C.D., and F. Rafi. 1978. Tanais stanfordi Richardson 1901 (Crustacea Tanaidacea) at the Fraser River estuary, British Columbia. Syesis, 11: 51 53.

Wu, R.S.S., and C.D. Levings. 1978. An energy budget for individual barnacles (Balanus glandula Darwin). Mar. Biol. (Berlin), 45: 225 235.

Wu, R.S.S., and C.D. Levings. 1979. Energy flow and population dynamics of the barnacle (Balanus glandula). Mar. Biol. (Berlin), 54: 83 89.

Levings, C.D. 1981. Consequences of training walls and jetties for aquatic habitats at two British Columbia estuaries. Coastal Engineering 4: 111-136.

Levings, C.D. 1980. Biology and energetics of Eogammarus confervicolus (Amphipoda, Anisogammaridae) at the Squamish River estuary. British Columbia. Can. J. Zool, 58: 1652 1663.

Pomeroy, W.M., and C.D. Levings. 1980. Association and feeding relationships between Eogammarus confervicolus (Stimpson) (Amphipoda, Gammaridae) and benthic algae on Sturgeon and Roberts Bank, Fraser River estuary, Can. J. Fish. Aquat. Sci., 37: 1 10.

Wu, R.S.S., and C.D. Levings. 1980. Mortality, growth and fecundity of transplanted mussel and barnacle populations near a pulp mill outfall. Mar. Poll. Bull. 11: 11 15.

Fournier, J.A., and C.D. Levings. 1982. Polychaetes recorded near two pulp mills on the north coast of British Columbia: a preliminary taxonomic and ecological account. Syllogeus (National Museum of Canada) 40: 1 91.

Levings, C.D. 1982. The ecological consequences of dredging and dredge spoil disposal in Canadian waters. Published by National Research Council of Canada Associate Committee on Scientific Criteria for Environmental Quality. NRCC Publication No. 18130. 141 p.

Levings, C.D., R.E. Foreman, and V.J. Tunnicliffe. 1983. A review of the benthos of the Strait of Georgia and contiguous fjords. Can. J. Fish. Aquat. Sci. 40: 1120 1141.

Seki, H., A. Otsuki, S. Daigobo, C.D. Levings, and C.D. McAllister. 1984. Microbial contribution to the mesotrophic ecosystem of the Campbell River estuary during summer. Archiv. fur Hydrobiologie 102: 215 228.

Levings, C.D. 1985. Juvenile salmonid use of habitats altered by a coal port in the Fraser River estuary, British Columbia. Mar. Poll. Bull. 17: 248 254.

Levings, C.D., E.P. Anderson and C.W. O’Connell. 1985. Biological effects of dredged material disposal in Alberni Inlet p. 132 155. In Ketchum, B., Capuzzo, J.M., Burt, W.V., Duedall, I.W., and P.K. Park. Wastes in the Ocean, Vol. 6: Nearshore Waste Disposal. John Wiley and Sons. 534 p.

Stanhope, M.J. and C.D. Levings. 1985. Growth and production of Eogammarus confervicolus (Amphipoda: Anisogammaridae) at a log storage site and in areas of undisturbed habitat within the Squamish estuary, British Columbia. Can. J. Fish. Aquat. Sci. 42: 1733 1740.

Levings, C.D., McAllister, C.D. and B.D. Chang. 1986. Differential use of the Campbell River estuary, British Columbia, by wild and hatchery reared juvenile chinook salmon (Oncorhynchus tshawytscha). Can. J. Fish. Aquat. Sci. 43: 1386 1397.

Swales, S., Lauzier, R.B., and C.D. Levings. 1986. Winter habitat preferences of juvenile salmonids in two interior rivers in British Columbia. Can. J. Zool. 64: 1506 1514.

Levings, C.D., C.D. McAllister, and J.S. Macdonald. 1987. Capacity of estuarine wetlands for juvenile salmonid production – Campbell River estuary, British Columbia, Canada. Northwest Environment Journal 3: 145 146.

Otsuki, A., H. Seki, C.D. McAllister, and C.D. Levings. 1987. Measurement of net growth rates of herbivorous benthic animals using periphyton labelled simultaneously with Carbon 13 and nitrogen 15 isotopes. Limnol. Oceangr. 32: 499 503.

Seki, Humitake, Akira Otsuki, Yoshiaki Hara, Kenneth V. Stephens, Colin D. Levings and Carey D. McAllister. 1987. Dynamics of organic materials in the Campbell River Estuary at the time of the spring bloom of phytoplankton. Arch. Hydrobiol. 111(2): 209 216.

Birtwell, I.K., C.D. Levings, J.S. Macdonald and I.H. Rogers. 1988. A review of fish habitat issues in the Fraser River system. Water Pollution Research Journal of Canada 23:1 30.

Macdonald, J.S., C.D.Levings, C.D. McAllister, U.H.M. Fagerlund, and J.R. McBride. 1988. A field experiment to test the importance of estuaries for Chinook salmon (Oncorhynchus tshawytscha) survival: short term results. Can. J. Fish. Aquat. Sci. 45: 1366 1377.

Rogers, I. H., J.A. Servizi and C.D. Levings. 1988. Bioconcentration of chlorophenols by juvenile chinook salmon (Oncorhynchus tshawytscha) overwintering in the upper Fraser River: field and laboratory tests. Water Poll. Res. J. Can. 23: 100 113.

Seki, H., A. Otsuki, Y. Fukuto, K.V. Stephens, C.D. Levings, H. Yamagishi, M. Kotyk and C.D. McAllister. 1988. Influence of wastewater discharge on the uptake kinetics of dissolved organic materials by microorganisms in the Fraser River estuary, British Columbia. Arch. Hydrobiol. 112(1): 55 65.

Swales, A., F. Caron, J.R. Irving and C.D. Levings. 1988. Overwintering habitats of coho salmon (Oncorhynchus kisutch) and other juvenile salmonids in the Keogh River system, British Columbia. Can. J. Zool. 66: 254 261.

Rogers, I.H., C.D. Levings, W.L. Lockhart, and R.J. Norstrom. 1989. Observations on overwintering juvenile chinook salmon (Oncorhynchus tshawytscha) exposed to bleached kraft mill effluent in the upper Fraser River, British Columbia. Chemosphere 19: 1853 1868.

Swales, S. and C.D. Levings. 1989. The role of off-channel ponds in the life-cycle of coho salmon (Oncorhynchus kisutch) and other juvenile salmonids in the Coldwater River, British Columbia. Can. J. Fish. Aquat. Sci. 46(2): 232 242.

Parsons, T.R., B.E. Rokeby, C.M. Lalli, and C.D. Levings. 1990. Experiments on the effect of salmon farm wastes on plankton ecology. Bull. Plankton Society of Japan 37:49-57.

Levings, C.D. and J.S. Macdonald. 1991. Rehabilitation of estuarine fish habitat at Campbell River, British Columbia. American Fisheries Society Symposium 10: 176-190.

Levings, C.D. 1991. Strategies for restoration and development of fish habitat in the Strait of Georgia – Puget Sound Inland Sea, northeast Pacific Ocean. Marine Pollution Bulletin 23: 417 – 422.

Levings, C.D, Conlin, K., and B. Raymond, 1991. Intertidal habitats used by juvenile chinook (Oncorhynchus tshawytscha) rearing in the North Arm of the Fraser River estuary Mar. Poll. Bull 22: 20-26.

Beamish, R.J. and C.D. Levings. 1991. Abundance and freshwater migrations of the anadromous parasitic lamprey (Lampetra tridentata) from a tributary of the Fraser River, British Columbia. Can. J. Fish. Aquat. Sci.48: 1250-1263.

Levings, C.D. and R.B. Lauzier. 1991. Extensive use of the Fraser River basin as winter habitat by juvenile chinook salmon (Oncorhynchus tshawytscha). Can. J. Zool. 69: 1759-1767.

Levings, C.D., Wells, P.G., and J.B. Sprague, 1992. Publications of Dr Michael Waldichuk . Mar Poll Bull 25: 1-13

Black, E.A. Gillis, D.J., Hay, D.E., Haegele, C.W., and C.D. Levings. 1992. Predation by caged salmon in British Columbia. Bull. Aquaculture Association of Canada 92-3: 58-60.

Levings, C.D., Hvidsten, N.A., and B.O. Johnsen. 1994. Feeding of Atlantic salmon (Salmo salar L) post-smolts in a fjord, central Norway. Can. J. Zoology 72: 834-839.

Levings, C.D. 1994. Some ecological concerns for net-pen culture of salmon on the coasts of the northeast Pacific and Atlantic Oceans, with special reference to British Columbia J. Applied Aquaculture 4: 65-141

Levings, C.D. 1994. Feeding behaviour of juvenile salmon and significance of habitat during estuary and early sea phase. Nordic Journal of Freshwater Research 69: 7-16.

Higgs, D.A., J.S. Macdonald, C.D. Levings, and B. Dosanjh. 1995. Nutrition and feeding habits of Pacific salmon (Oncorhynchus spp.) in relation to life history stage p.161-315. In Physiological ecology of Pacific salmon. Edited by Brett, R., Clarke, W.C., Groot, K., and L. Margolis. University of B.C. Press, Vancouver, B.C. 510 p.

Levings, C.D., D.E. Boyle and T.R. Whitehouse. 1995. Distribution and feeding of juvenile salmon in freshwater tidal creeks, lower Fraser River, British Columbia. Fisheries Management and Ecology. 2: 299-308

Levings, C.D., A. Ervik, P. Johannessen and J. Aure, 1995. Ecological criteria used to site fish farms in fjords. Estuaries 18-1A: 81-90.

Richardson, J.S. and C.D. Levings. 1996. Chlorinated organic contaminants in benthic organisms of the lower Fraser River, British Columbia. Water Pollution. Res. J. Canada. 31: 153-162

Gregory, R.S. and C.D. Levings. 1996. The effects of turbidity and vegetation on the risk of juvenile salmon (Oncorhynchus spp.) to predation by adult trout (O. clarkii). Environmental Biology of Fishes 47(3): 279-288

Grout, J., Levings, C.D. and J.S. Richardson, 1997. Decomposition rates of purple loosestrife (Lythrum salicaria) and Lyngbyei’s sedge (Carex lyngbyei) in the Fraser River estuary. Estuaries 20:96-102

Levings, C.D. and D.J.H. Nishimura, 1997. Created and restored marshes in the lower Fraser River, British Columbia: a summary of their functioning as fish habitat. Water Quality Research Journal of Canada 32:599-618

Gregory, R.S. and C.D. Levings 1998. Turbidity reduces predation on migrating juvenile Pacific salmon. Trans Amer Fish Soc 127: 275-285

Mjaavatten, O., Levings, C.D., and P. Poon 1998 Variation in the fatty acid composition of juvenile Chinook and coho salmon from the Fraser River estuary determined by multivariate analysis: role of environment and genetic origin. Comparative Biochemistry and Physiology 120B(2):291-309

Levings, C.D. 1998. The Fraser River estuary, British Columbia, Canada: an introduction to its ecology, habitat management systems, and possible comparative work at the Han River estuary, Korea. Ocean Research (Korea) 20(2):199-207

Levings, Colin D. 1999. An overview assessment of compensation and mitigation techniques used to assist fish habitat management in British Columbia estuaries. In Sustainable fisheries management: Pacific salmon. Edited by E. Eric Knudsen, Cleveland R. Steward, Donald D. MacDonald, Jack E. Williams, and Dudley W. Reiser. CRC Lewis Publishers, Boca Raton, New York. pp. 341-347.

Powles, H., M Bradford, R. Bradford, W. Doubleday, S. Innes, C.D. Levings. 2000 Assessing and Protecting Endangered Marine Species. ICES Journal Marine Science 57: 669-676

Emmett, R.L., A. Copping, P. Fishman, M. Hornberger, C.D. Levings, J. Newton, and R. Thom. 2000 Geographic signatures of West Coast estuaries. Estuaries 23: 765-792

Barry, K.L., Grout, J.A., Levings, C.D., Nidle, B., and G.E. Piercey, 2000. Impacts of acid mine drainage on juvenile salmonids in an estuary near Britannia Beach in Howe Sound, British Columbia Can J Fish Aquat Sci 57:2032-2043

Jamieson, G.S. and C.D. Levings, 2001. Marine protected areas in Canada – implications for both conservation and fisheries management Can J Fish Aquat Sci 58: 1-19

Sutherland, T.F., A.J. Martin, and C.D. Levings 2001. The characterization of suspended particulate matter surrounding a salmonid net-pen in the Broughton Archipelago, British Columbia. ICES Journal of Marine Science 58:404-410

Sutherland, T.F., Levings, C.D., Elliott, C.C. and W.W. Hesse, 2001. The effect of a ballast water treatment system on the survivorship of natural populations of marine plankton. Marine Ecology (Progress Series) 210: 139-148

Grout, J.A. and C.D. Levings, 2001. Effects of acid mine drainage from an abandoned copper mine, Britannia Mines, Howe Sound, BC, on transplanted blue mussels (Mytilus edulis)
Marine Env Res 51: 265-288

Jamieson, G.S., Foreman, M., Cherniawsky, J, and C.D. Levings, 2002. European Green Crab (Carcinus maenas) Dispersal: The Pacific Experience p. 561-578 in Paul, A.J., Dawe, E.G., Elner, R., Jamieson, G.S., Kruse, G.H., Otto, R.S., Sainte-Marie, B., Shirley, T.C, and D.Woodby (Eds). Crabs in Cold Water Regions: Biology, Management, and Economics. University of Alaska Sea Grant, AK-SG-02-01. Fairbanks, Alaska.

Arvai , J.L.,C.D. Levings, C.D., P.J. Harrison, and W.E. Neill, 2002. Improvement of the sediment ecosystem following diversion of an intertidal sewage outfall at the Fraser River estuary, Canada, with emphasis on Corophium salmonis (Amphipoda). Marine Pollution Bulletin 44: 511-519

Levings, C.D., Kieser, D., Jamieson, G.S., and S. Dudas.2002 .Marine and estuarine alien species in the Strait of Georgia, British Columbia. P. 111-131 In Claudia, Renata, Nantel, Patrick and Elizabeth Muckle-Jeffs (Eds). Alien invaders in Canada’s Waters, Wetlands, and Forests. Natural Resources Canada, Canadian Forest Service, Ottawa

Si-Wan Lee, Hansoo Lee Jeong-Chil Yoo, Jong-Geel Je, Colin Levings, and Woon-Kee Paek, 2002. Factors affecting the Conservation and Distribution of Migratory Waterbirds in the Southern Tidal Flats of Ganghwa Island. Korea Korean Journal of Environment and Ecology 16: 34-45 (in Korean)

Marsden, A.D. Robert E. DeWreede, and Colin D. Levings, 2003. Survivorship and growth of Fucus gardneri after transplant to an acid mine drainage-polluted area. Marine Pollution Bulletin 46:65-73

Levings, C.D., Stein, J.E., Stehr, C.M., and S.C. Samis, 2004. Introduction to the PICES practical workshop: objectives, overview of the study area, and projects conducted by the participants. Marine Environmental Research 57: 3-18

Levings, C.D. and S. M. Ong, 2004. Fish communities and life history attributes of English sole (Pleuronectes vetulus) in Vancouver Harbour. Marine Environmental Research 57:103-120

Larson, M., Foreman, M.G.G., Levings, C.D., and M.R. Tarbotton, 2004. Dispersion of discharged ship ballast water in Vancouver harbour, Juan de Fuca Strait, and offshore of the Washington coast. J Environmental Engineering and Science 2: 163-176

Horiguchi, T., Z. Li, S. Uno, M. Shimizu, H. Shiraishi, M. Morita, J. A. J. Thompson and C. D. Levings, 2004. Contamination of organotin compounds and imposex in molluscs from Vancouver, Canada. Marine Environmental Research 57:75-88

Je, Jong-Geel,Tatyana Belan, Colin Levings and Bon Joo Koo, 2004. Changes in benthic communities along a presumed pollution gradient in Vancouver Harbour
Marine Environmental Research 57: 121-135

Romanuk, T.N. and C.D. Levings, 2003. Associations between arthropods and supralittoral ecotone: dependence of aquatic and terrestrial taxa on riparian vegetation. Environmental Entomology 32:1343-1353

Sutherland, T., Levings, C.D., Petersen, S. and W. Hesse, 2003. Mortality of zooplankton and invertebrate larvae exposed to cyclonic pre-treatment and ultraviolet radiation. Marine Technology Society Journal 37: 3-12

Troffe, P.M., Ong, S., Levings, C.D. and T.F. Sutherland, 2003. Anatomical damage to humpback shrimp (Pandalus hypsinotus) (Brandt 1851) caught by trawling and trapping. J. Shellfish Research 22: 561-568

Levings, C.D., Cordell, J.R., Ong, S, and G.E. Piercey, 2004. The origin and identity of invertebrate organisms being transported to Canada’s Pacific coast by ballast water. Can J Fish Aquat Sci 61:1-11

Levings, C.D., K.L. Barry, J.A. Grout, G.E. Piercey, A.D. Marsden, A. Coombs, and B. Mossop. 2004. Effects of acid mine drainage on the estuarine food web, Britannia Beach, Howe Sound, British Columbia, Canada. Hydrobiologia 525: 185-202

Levings, C.D. and T.G. Northcote, 2004. Effects of forestry on estuarine ecosystems supporting fishes. 320-335 Chapter 15 in Fishes & Forestry – Worldwide Watershed Interactions & Management. T.G. Northcote and G.F. Hartman [editors] Blackwell Science, Oxford

Bendell-Young, L., Yin, Kedong, Thomas, C., Harrison, P.J., Feeney, T., Arvai, J.L., Levings, C.D., and L. Ross, 2004. Biogeochemistry of the intertidal area of the Fraser River estuary in B.J. Groulx, Mosher, D.C., Luternauer, JL. and D. E. Bilderback (Ed). Fraser River Delta, British Columbia: Issues of an Urban Estuary Geological Survey of Canada Bulletin 567:189-212

Levings, C.D. 2004. Knowledge of fish ecology and its application to habitat management in B.J. Groulx, Mosher, D.C., Luternauer, JL. and D. E. Bilderback (Ed). Fraser River Delta, British Columbia: Issues of an Urban Estuary. Geological Survey of Canada Bulletin 546: 213-236

Kang, Rae-Seon, Heung-Sik Park, Ki-Sik Won, Jong-Man Kim and Colin Levings, 2005. Competition as a determinant of the upper limit of subtidal kelp Ecklonia stolonifera Okamura in the southern coast of Korea. J. Exp Marine Biol Ecology 314:41-52

Romanuk, T.N. and Colin D. Levings, 2005. Stable isotope analysis of trophic position and terrestrial versus marine carbon sources for juvenile Pacific salmonids in nearshore marine habitats. Fisheries Management and Ecology 11: 113-121

Yeats, P.A. Milligan, T.G.,T.F. Sutherland, T.F., Robinson, S.M.C., Smith, J.A.,. Lawton, P., and C.D. Levings, 2005. Lithium-Normalized Zinc and Copper Concentrations in Sediments as Measures of Trace Metal Enrichment due to Salmon Aquaculture. Chapter in Hargrave, B (Ed). Environmental Effects of Marine Finfish Aquaculture. Handbook of Environmental Chemistry, Vol 5: Water Pollution, Part M. Springer-Verlag. (published on line July 15 2005)

Stucchi, D., Terri-Ann Sutherland, Higgs, D., and C.D. Levings (2005) Near-Field Depositional Model for Finfish Aquaculture Waste. Chapter in Hargrave, B (Ed). Environmental Effects of Marine Finfish Aquaculture. Handbook of Environmental Chemistry, Vol 5: Water Pollution, Part M. Springer-Verlag. (published on line July 15 2005)

Troffe, P.M., Levings, C.D., Piercey, G.E., and V. Keong, 2005.
Preliminary observations on ecology and the effects of beam trawling and trapping on the sea whip (Halipteris willemoesi (Cnidaria: Octocorallia: Pennatulacea)) in British Columbia, Canada. Aquatic Conservation: Marine and Freshwater Ecosystems 15: 523-533

Romanuk, T.N. and C. D. Levings 2006.Relationships between fish and supralittoral vegetation in nearshore marine habitats. Aquatic Conservation: Marine and Freshwater Ecosystems 16:115-132

Levings, C.D. , D.E. Varela, N.M. Mehlenbacher, K.L. Barry, G.E. Piercey, M. Guo and P.J. Harrison, 2005. Effect of an acid mine drainage effluent on phytoplankton biomass and primary production at Britannia Beach, Howe Sound, British Columbia. Marine Pollution Bulletin 50: 1585-1594

Nelson, W.A, Levings, C.D., and A.J. Paul (2007) Population Growth and Critical Demography of White Sturgeon in the Lower Fraser River. . American Fisheries Society Symposium 56:381-396

Lee, C.G. and C. D. Levings, 2007. Note: The Effects of Temperature and Desiccation on Surf Smelt (Hypomesus pretiosus) Embryo Development and Hatching Success: Preliminary Field and Laboratory Observations. Northwest Science 81: 166-171

Sutherland, T.A., Petersen, S, Levings, C.D. and A. Martin, 2007. Distinguishing between natural and aquaculture-derived sediment concentrations of heavy metals in the Broughton Archipelago, British Columbia. Marine Pollution Bulletin 54: 1451-1460

Sutherland, T.F., CD Levings, SA Petersen, P Poon, B . Piercey, 2007. The use of meiofauna as an indicator of benthic organic enrichment associated with salmonid aquaculture Marine Pollution Bulletin, 2007 54:1249-1261

Sutherland, T.A., Galloway, J., Hare, R., and C.D. Levings. 2007 The calibration of multibeam acoustic technology (EM3000) with sediment properties. Estuaries and Coasts 75:447-458

Romanuk, T.N. and C.D. Levings, 2010. Reciprocal Subsidies and Food Web Pathways Leading to Chum Salmon Fry in a Temperate Marine-Terrestrial Ecotone. PLosONE 5(4):e10073

DiBacco, C., Humphrey, D.B., Nasmith, L.E., and C. D. Levings, 2011. Ballast Water Transport of Non-Indigenous Zooplankton to Canadian Ports. ICES Journal of Marine Science 69: 483-491

Mach, M.E., Levings, C.D., Chan, K.M.A., and P.S. McDonald, 2012.An Atlantic Infaunal Engineer is Established in the Northeast Pacific: Clymenella torquata (Polychaeta: Maldanidae) on the British Columbia and Washington Coasts. Biological Invasions.14(3):503-507

Lo, V. B., Levings, C.D., and K. M. A. Chan, 2011.Quantifying potential propagule pressure of aquatic invasive species from the commercial shipping industry in Canada. Marine Pollution Bulletin 64: 295-302

Briski, E., Bailey, S., Casas-Monroy, O., DiBacco, C., Kaczmarska, I., Levings, C., MacGillivary, M. McKindsey, C., Nasmith, L., Parenteau, M., Piercey, G., Rochon, R., Roy, S., Simard, N., Villac, M., Weise, A., MacIsaac, H.2012. Relationship between propagule pressure and colonization pressure in invasion ecology. Proceedings of the Royal Society of London B: Biological Sciences, rspb20112671

Naiman, Robert J., J. Richard Alldredge, David A. Beauchamp Peter A. Bisson, James Congleton, Charles J. Henny, Nancy Huntly, Roland Lamberson ,Colin Levings, Erik N. Merrilll, William G. Pearcy, Bruce E. Rieman, Gregory T. Ruggerone, Dennis Scarnecchia, Peter E. Smouse, and Chris C. Wood, 2012 Developing a broader scientific foundation for river restoration: Columbia River food webs. Proceedings of the National Academy of Sciences 109, no. 52 (2012): 21201-21207.

Sutherland, T.F and C.D. Levings, 2013. Quantifying Non-indigenous Species in Accumulated Ballast Slurry Residuals (Swish) Arriving at Vancouver, British Columbia. Progress in Oceanography 115:211-218

Briski, E., Bailey, S.,Casas-Monroy, O., DiBacco, C., Kaczmarska, I., Levings, C., MacGillivary, M. McKindsey, C., Nasmith, L., Parenteau, M., Piercey, G., Rochon, R., Roy, S., Simard, N.,Villac, M., Weise, A., MacIsaac, H.2013 Taxon- and vector-specific variation in species richness and abundance during the transport stage of biological invasions. Limnology and Oceanography 58:1361-1372

Rieman, B. E., Smith, C. L., Naiman, R. J., Ruggerone, G. T., Wood, C. C., Huntly, N., Levings, C.D…. & P. Smouse, P. (2015). A comprehensive approach for habitat restoration in the Columbia basin. Fisheries, 40(3), 124-135.

Levings, C.D., 2015. Estuary conservation. Entry in Kennish, M. (Ed). Encyclopedia of Estuaries. Springer.

Levings, C.D., 2015. Estuary sustainability. Entry in Kennish, M. (Ed). Encyclopedia of Estuaries. Springer.

Levings, C.D., 2015. Estuary conservation zone. Entry in Kennish, M. (Ed). Encyclopedia of Estuaries. Springer.

Mach, M.E., Levings, C.D., and Kai Chan, 2017. Nonnative species in British Columbia eelgrass beds spread via shellfish aquaculture and stay for the mild climate. Estuaries and Coasts 40(1):187-199

Levings, C.D.. 2017. Salmon. In: Bodtker, K. (Ed.). 2017. Ocean Watch: Howe Sound Edition. Coastal Ocean Research Institute, Vancouver Aquarium Marine Science Centre. Vancouver, B.C., Canada. 365 pp. Available online: http://oceanwatch.ca

Levings, C.D. 2020 Edward Flanders Ricketts and Marine Ecology of the Inner Coast of British Columbia. Archives of Natural History 47 (2):115-123

Levings, C. 2020. Connecting to Edward Flanders Ricketts Through Mentors and an Iconic Ship Sailing in the Gulf of Alaska. Steinbeck Review 17 (1):73-81

Levings, C.2021. The Fishery Technology Complex- From Mapping to Depletion of Pacific Ocean Perch 1880s-1970s. Technology and Culture 62 (1): 185-211

Chalifour, L., Holt, C., Camaclang, A. E., Bradford, M. J., Dixon, R., Finn, R. J. R., Hemming, V., Hinch, S. G., Levings, C. D., MacDuffee, M., Nishimura, D. J. H., Pearson, M., Reynolds, J. D., Scott, D. C., Spremberg, U., Stark, S., Stevens, J., Baum, J. K., & Martin, T. G. 2022. Identifying a pathway towards recovery for depleted wild Pacific salmon populations in a large watershed under multiple stressors. Journal of Applied Ecology, 59, 2212– 2226. https://doi.org/10.1111/1365-2664.14239

Levings, C.D. and C.R. Bates, 2022. The intertidal fish collections of Ed Ricketts at Tofino, British Columbia, Canada, 1945 and 1946. Canadian Field Naturalist 136(2): 133-137